| 1987 | B.S.; University of Tsukuba (Biological Sciences) |
| 1989 | M.S.; University of Tsukuba (Biological Sciences) |
| 1993 | Ph.D.; University of Tsukuba (Biological Sciences; Biochemistry and Biophysics) |
| 1992-1994 | JSPS Research Fellow;The University of Tokyo |
| 1994-1997 | JSPS Postdoctoral Fellow for Research Abroad and Brown Postdoctoral Fellow; Yale University, USA |
| 1997-1999 | Postdoctoral Fellow; University of Michigan, USA |
| 1999-2005 | Assistant Professor; University of Washington, USA |
| 2005-2009 | Associate Professor; University of Washington, USA |
| 2008-2012 | JST PRESTO Researcher |
| 2009-present | Professor, University of Washington; USA |
| 2011-present | Distinguished Professor of Biology; University of Washington, USA |
| 2011-present | Investigator; Howard Hughes Medical Institute and Gordon and Betty Moore Foundation (HHMI-GBMF), USA |
| 2013-present | Visiting Professor; WPI-ITbM, Nagoya University |